Partial audio deepfakes manipulate only selected speech regions, making them difficult to be localized. Existing methods exploit boundary cues for partial deepfake localization, but primarily focus on identifying boundary positions rather than modeling the feature changes that characterize authenticity transitions. Meanwhile, the internal characteristics of continuous bona fide and spoofed segments remain underexplored. In this paper, we propose Boundary and Intra-Segment Learning (BISL), which introduces boundary learning to model feature differences between adjacent frames and distinguish authenticity transitions from general acoustic variations. In addition, intra-segment learning captures the overall characteristics of continuous bona fide and spoofed segments while enhancing feature consistency within each segment. By jointly learning frame, boundary, and segment information, BISL enables more effective fine-grained partial audio deepfake localization. Experiments on multiple localization benchmarks show that BISL achieves an EER of 2.52\% and an F1-score of 97.40\% on PartialSpoof, outperforming the compared methods, while maintaining competitive performance on HAD and improved cross-dataset performance on LPS. The code will be made publicly available upon acceptance.

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