Non-fixed flexible antenna architectures, such as fluid antenna system (FAS), movable antenna (MA), and pinching antenna, have garnered significant interest in recent years. Among them, rotatable antenna (RA) has emerged as a promising technology for enhancing wireless communication and sensing performance through flexible antenna orientation/boresight rotation. By enabling mechanical or electronic boresight adjustment without altering physical antenna positions, RA introduces additional spatial degrees of freedom (DoFs) beyond conventional beamforming. In this paper, we provide a comprehensive tutorial on the fundamentals, architectures, and applications of RA-empowered wireless networks. Specifically, we begin by reviewing the historical evolution of RA-related technologies and clarifying the distinctive role of RA among flexible antenna architectures. Then, we establish a unified mathematical framework for RA-enabled systems, including general antenna/array rotation models, as well as channel models that cover near- and far-field propagation characteristics, wideband frequency selectivity, and polarization effects. Building upon this foundation, we investigate antenna/array rotation optimization in representative communication and sensing scenarios. Furthermore, we examine RA channel estimation/acquisition strategies encompassing orientation scheduling mechanisms and signal processing methods that exploit multi-view channel observations. Beyond theoretical modeling and algorithmic design, we discuss practical RA configurations and deployment strategies. We also present recent RA prototypes and experimental results that validate the practical performance gains enabled by antenna rotation. Finally, we highlight promising extensions of RA to emerging wireless paradigms and outline open challenges to inspire future research.

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