We study the fine-grained complexity of evaluating Boolean bounded-variable first-order queries over sparse relational structures. For every fixed $k \ge 2$, every relational signature, and every fragment between $k$-variable primitive positive ($\mathrm{PP}^{k}$) and first-order ($\mathrm{FO}^{k}$) logic, we prove, assuming the Sparse MAX-$3$-SAT hypothesis, a dichotomy theorem for evaluation in $O(m^{k-\varepsilon})$ time, where $m$ is the number of tuples in the input structure. The only tractable cases fall into three families: (1) three-variable fragments, (2) two-variable fragments, and (3) fragments over unary signatures. On the hard side, for every fixed $k \ge 4$ and every $\varepsilon > 0$, there is a fixed sentence $\varphi_\varepsilon$ in $\mathrm{PP}^{k}$ over a single binary relation symbol, depending on $\varepsilon$ but not on the input structure, whose evaluation cannot be performed in $O(m^{k-\varepsilon})$ time. For the tractable cases, we show that the evaluation problem can be solved in $2^{O(|\varphi|)} \cdot m^{k-\varepsilon}$ time for some $\varepsilon > 0$. Moreover, every tractable fragment reduces, with a $2^{O(|\varphi|)}$ blowup in formula size under DAG representations, to one of the following query languages: (1) Tarski's calculus of relations, (2) a new Boolean modal logic for sparse model checking, and (3) one-variable counting logic.

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