A connectome-constrained model of the fly visual system, optimized for motion and then frozen, can be driven over architectural drawings by prescribed motion and used as a texture representation. We compare it with information-matched baselines that see the same 721 photoreceptor samples. On clean synthetic data the frozen model transfers but loses to task training: 0.857 area-weighted accuracy in one-shot hatch matching versus 0.959 for a 5,888-parameter CNN, and 0.619 IoU in wall segmentation versus 0.905 for a matched network. Under scan noise and thickened strokes, the trained networks lose up to 0.188 accuracy while the frozen pipeline loses 0.030. On fourteen production sheets, opened once, a 1,876-parameter fly model reaches 0.505 average precision versus 0.415 for a network two hundred times larger. A preregistered held-out split confirms the clean-data ordering: 0.835 for the circuit, 0.894 for receptors only, and 0.971-0.980 for trained CNNs. Rewiring the connectome while preserving degrees or type pairs and transmitter signs costs 0.271-0.356 accuracy across three seeds, so the exact wiring is load-bearing. Yet the intact circuit does not beat its moving retina, and T4/T5 silencing leaves both tasks intact. Longer observations reverse the circuit-receptor ordering once the stimulus spans a period, but not through T4/T5. Thus active sampling and exact structure matter, while clean-data practical performance remains dominated by task-trained networks and the useful transfer margin is largely retinal.

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