Large language models (LLMs) have demonstrated growing competence in generating web pages from UI screenshots, which convey both visual structure and cues to application behavior. Yet most screenshot-to-code benchmarks emphasize visual fidelity, while interactive generation benchmarks often supply behavioral specifications or demonstrated transitions. Whether models can infer and realize interactions from static screenshots alone remains insufficiently evaluated. We introduce UI2App to evaluate interaction inference: inferring and realizing application behavior from static visual cues without added behavioral guidance. UI2App comprises 600 screenshots organized into 95 state-coherent sets for runnable multi-route web applications. Our end-to-end pipeline evaluates each artifact along three dimensions: executability, visual fidelity, and interaction inference. The interaction metric (IIS) assesses functional correctness and state-management complexity, crediting valid implementations rather than requiring a match to a single reference. Experiments on six frontier vision-language models reveal a marked mismatch between visual fidelity and interaction realization: the visual-fidelity leader scores only 8.1 on IIS, ranking fourth, while the IIS leader achieves 4.5 times that score. High-complexity interactions such as cross-route state persistence remain a major bottleneck, with five of the six models scoring at most 3.5 on this dimension. Overall, these results highlight interaction inference as a key challenge in generating functional web applications from static screenshots.

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