Software engineering has long inferred where a system's knowledge resides from who authored its code. The truck factor, the Degree-of-Authorship metric, and the degree-of-knowledge model all rest on one inference -- that authoring a region of code is evidence of understanding it -- and for most of software's history it was a workable proxy, because code entered a repository only when a human wrote it, which forced at least transient understanding. This paper argues that AI code generation severs that inference at its root, and that the consequence is not the degradation of the authorship-based metrics but their invalidation as a class. When an agent generates a module and a human merges it, the version-control record still attributes authorship, but the attribution no longer licenses any conclusion about comprehension: the same footprint is now compatible with full, partial, or no understanding. The metric still returns a number; the number measures a substrate that has come uncoupled from the quantity it was used to estimate. The collapse is corroborated by the field's own measurement failures, and the methodological corollary is load-bearing: the instrument the comprehension-debt era needs cannot be built by refining the knowledge-concentration metrics, because no function of an authorship footprint recovers an inference the footprint no longer supports. The replacement must be grounded in evidence of comprehension rather than authorship. I state a falsifiable prediction that discriminates the two -- that systems with a healthy authorship-derived truck factor but low comprehension-measured retention will suffer incident-resolution failures the authorship metric does not predict -- and argue that building the comprehension-grounded instrument at the scale of a system and a team is the field's open measurement problem, left open here.

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